Main Focus

  • Respiration in trees and soils (rates, substrates, processes reflected in geochemical proxies)
  • Carbon allocation (use of old nonstructural carbon, fine roots processes)
  • Soil organic matter

Curriculum Vitae

2008 - 2011    B.Sc in Environmental Science, Geography, and division of Geoinformatics, with honor. The Hebrew University of Jerusalem

2011 - 2018    Ph.D. in Environmental Science. The Hebrew University of Jerusalem, under the supervision of Prof. Alon Angert. Thesis: “Better Understanding of Tree Stem Respiration by Combining O2, CO2, and O2 Isotopes Measurements”

2018 - present   Postdoctoral position, Max Planck Institute for Biogeochemistry.


Hilman, B., Solly, E. F., Kuhlmann, I., Brunner, I., and F. Hagedorn (2024), Species-specific reliance of trees on ectomycorrhizal fungi for nitrogen supply at an alpine treeline. Fungal Ecology 71, 101361.

Rog, I., Hilman, B., Fox, H., Yalin, D., Qubaja, R., and T. Klein (2024), Increased belowground tree carbon allocation in a mature mixed forest in a dry versus a wet year. Global Change Biology 30 (2), e17172 (2024)

Salomón, R. L., Helm, J., Gessler, A., Grams, T. E. E., Hilman, B., Muhr, J., Steppe, K., Wittmann, C., and H. Hartmann (2024), The quandary of sources and sinks of CO2 efflux in tree stems - new insights and future directions. Tree Physiology 44 (1).

Helm, J.Muhr, J.Hilman, B., Kahmen, A., Schulze, E. D.Trumbore, S. E.Herrera-Ramirez, D., and H. Hartmann (2023), Carbon dynamics in long-term starving poplar trees—the importance of older carbohydrates and a shift to lipids during survival. Tree Physiology, tpad135.

Helm, J., Salomón, R. L., Hilman, B.Muhr, J., Knohl, A., Steppe, K., Gibon, Y., Cassan, C., and H. Hartmann (2023), Differences between tree stem CO2 efflux and O2 influx rates cannot be explained by internal CO2 transport or storage in large beech trees. Plant, Cell and Environment 46 (9), pp. 2680 - 2693.

Hilman, B., Weiner, T., Haran, T., Masiello, C. A., Gao, X., and A. Angert (2022), The apparent respiratory quotient of soils and tree stems and the processes that control it. Journal of Geophysical Research: Biogeosciences 127 (3), e2021JG006676.

Hilman, B., J. Muhr, J. Helm, I. Kuhlmann, E.-D. Schulze, and S. Trumbore (2021), The size and the age of the metabolically active carbon in tree roots, Plant, Cell & Environment44(8), 2522-2535,

Helm, J., H. Hartmann, M. Göbel, Hilman, B., D. A. Herrera, and J. Muhr (2021), Low-cost chamber design for simultaneous CO2 and O2 flux measurements between tree stems and the atmosphere, Tree Physiology 41 (9), 1767-1780,

Hilman, B., T. Weiner, T. Haran, and A. Angert (2020), Studying Soil and Tree Stem Respiration in Mediterranean oak forest using the Respiratory Quotient, edited, American Geophysical Union, Washington,

Hicks Pries, C., A. Angert, C. Castanha, Hilman, B., and M. S. Torn (2020), Using respiration quotients to track changing sources of soil respiration seasonally and with experimental warming, Biogeosciences17(12), 3045-3055, https://10.5194/bg-17-3045-2020.

Hilman, B., et al. (2019), Comparison of CO2 and O2 fluxes demonstrate retention of respired CO2 in tree stems from a range of tree species, Biogeosciences16(1), 177-191, https://10.5194/bg-16-177-2019.

Weiner, T., A. Gross, G. Moreno, M. Migliavacca, M. Schrumpf, M. Reichstein, Hilman, B. A. Carrara, and A. Angert (2018), Following the Turnover of Soil Bioavailable Phosphate in Mediterranean Savanna by Oxygen Stable Isotopes, Journal of Geophysical Research: Biogeosciences123(6), 1850-1862,

Hilman, B., and A. Angert (2018), Discrimination in Tree Stems O2 Uptake and the Dole Effect, Global Biogeochemical Cycles32(8), 1208-1225, https://10.1029/2018gb005918.

Hilman, B., and A. Angert (2016), Measuring the ratio of CO2 efflux to O2 influx in tree stem respiration, Tree Physiol36(11), 1422-1431, https://10.1093/treephys/tpw057.

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